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Whole-Grain Spaghetti with Veggi-fied Meat Sauce

Ingredients

• 6 ounces whole-grain spaghetti (such as Barilla)
• Cooking spray
• 1 cup finely chopped onion
• 8 ounces 93% lean ground beef
• 6 garlic cloves, minced
• 1 cup finely chopped zucchini
• 8 ounces cremini mushrooms, finely chopped
• 1 (14.5-ounce) can unsalted diced tomatoes
• 1 tablespoon unsalted tomato paste
• 1/2 teaspoon dried oregano
• 1/4 teaspoon crushed red pepper
• 3/4 teaspoon kosher salt
• 1/4 teaspoon freshly ground black pepper
• 2 tablespoons finely grated Parmigiano-Reggiano cheese

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